Fundamental tissue characteristics and functions

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Simon Doyle
Fundamental tissue characteristics and functions

The fundamental tissue or soil tissue, in botany, is a tissue composed of cells of the parenchyma (mainly), collenchyma and sclerenchyma. The cells of these tissues can be located throughout the plant or in specific places or structures, have different morphological characteristics and perform multiple functions in the plant.

The functions of this tissue are essential for the survival of the plant, since it participates in storage, structural and mechanical support, food production (via photosynthesis), regeneration, among other functions..

Parenchyma and other tissue structures of a plant. Taken and edited from: Marion Moseby [CC BY-SA 4.0 (https://creativecommons.org/licenses/by-sa/4.0)].

Article index

  • 1 Features
    • 1.1 -Parenchyma cells
    • 1.2 - Collenchyma cells
    • 1.3 - Sclerenchyma cells
  • 2 Functions
    • 2.1 Parenchyma cells
    • 2.2 Collenchymal cells
    • 2.3 Sclerenchyma cells
  • 3 References

Characteristics

The fundamental tissue is made up of three types of cells:

-Parenchyma cells

They are the most abundant cells in the fundamental tissue that come from the parenchymal tissue, a poorly specialized tissue formed by living cells. These cells have a complex physiology, have vacuoles and their primary walls are thin, although on rare occasions they can become thick..

Additionally, these cells divide by mitosis and remain alive after reaching maturity. They have various shapes that will depend on their location in the plant as well as their function; These shapes can be imperfect spherical, stellate, polyhedral and even branched.

They have air-filled spaces at the vertices or corners of the cell. They generally do not have chloroplasts (with some exceptions), but they do have leukoplasts. Its vacuoles store tannins, crystals and other compounds.

Cell types

Chlorophyll

Cylindrical cells perpendicular to the surface, which present abundant chloroplasts and are separated by intercellular spaces. They are found in the green areas of the plant, under the epidermis.

Cells form two types of chlorophyll tissue; the so-called lagoon or spongy tissue, which is located in the part where there is more shadow on the leaf, and the palisade tissue, located in the area where there is greater exposure to sunlight.

Bookers

Cells without chloroplasts are abundant in rhizomes, aerial stems, and in roots such as potatoes, beets, and carrots. They have also been observed in seeds, fruit pulps and in the stem of sugar cane..

Aeriferous

They are typical plant cells that inhabit aquatic and humid environments. They have irregular shapes, with large spaces between one cell and another. They are found in both roots and stems.

At least three production mechanisms are known for these cells and tissues, which are related to the way in which gas spaces or cavities are created..

  • Schizogeny: the formation of air spaces occurs by cell differentiation, during the development of the organ.
  • Lysogenesis: occurs under environmental stress and gaseous spaces are formed by cell death.
  • Expansigenia: this last mechanism is not recognized by the entire botanist community, however it is thought that it occurs without the need for cell junctions to disappear.
Aquifers

They are cells that store water. Although almost all cells do, in these the proportions of the liquid are higher than in the rest, that is, they have a high degree of specificity for this function. They are large, vacuolated cells, with thin walls. They are located in the underground organs.

They are characteristic of xerophilous plants (cacti and prickly pears for example), that is, they live in dry environments.

-Collenchyma cells

They are responsible for giving elasticity and solidity to the plant, they are living cells. These cells agglomerate or form a compact mass, they remain alive after maturing. They have walls composed of pectin and cellulose, with secondary thickening or widening of irregular shape. They do not have lignin.

They have a rectangular, elongated or prismatic shape, that is, in the shape of a polyhedron. When they are cross-cut, they are polygonal. They can measure up to 2 millimeters and generally do not present chloroplasts, but sometimes they present tannins..

Cell types

Angular

Cells whose walls have a pronounced thickening at the angles where they join with other cells.

Tangential

Cells that have thickening in the walls that are parallel (tangential) to the surface of the organ.

Lagoons

The cells present thickening or widening of the walls towards the intercellular spaces.

Celery Colenchyme (Apium graveolens). Taken and edited from: Sahaquiel9102 [CC BY 3.0 (https://creativecommons.org/licenses/by/3.0)].

-Sclerenchyma cells

They are dead cells, they have a thickened secondary wall, composed of cellulose, hemicellulose and lignin. They die upon reaching maturity. They are arranged in a compact mass.

Cell types

Sclerenchymal fibers

They come in a wide variety of shapes and sizes. They are classified according to their location in the plant. They have secondary walls with lignin. Sometimes they are nucleated living cells.

Stony

Also called sclereids, they present a great variety of forms; they can be short, elongated, with thin and bulging shapes distally, polyhedral, branched, etc. They are generally dead cells, with walls that vary in thickness. They are found throughout the body of the plant.

Features

As we have already described before, the fundamental tissue or system is made up of cells from three different tissues and their functions are as follows:

Parenchyma cells

These cells have multiple functions in the plant. In the first place, its function is to reactivate meristematic activity, an activity responsible for plant growth. These cells are responsible for tissue regeneration, healing, and the production of new roots and shoots..

They participate in photosynthesis, food production, and gas exchange; they also store sugars, fats, proteins, and water. They are part of the filler tissue of any organ of the plant and also provide buoyancy to some aquatic plants.

Collenchyma cells

The cells that form the collenchyma tissue are responsible for providing support and structure to the plants, essentially in the growth areas, such as leaves and shoots, not in roots. They also provide support and support in adult plant organs that do not produce much sclerenchyma..

Sclerenchyma cells

These cells, like those of the collenchyma, make up the tissue that provides support and support to the plant that has stopped lengthening or growing. It provides the plant with elasticity and resistance to mechanical actions such as twisting, weight or stretching..

The presence of lignin and the thick and hard walls in these cells are the basis of the strength and rigidity of the cell, and also protect it from external physical, biological and chemical attacks..

Cross section of Sansevieria sp. The epidermis, chlorenchyma, and sclerenchyma fiber are observed. Taken and edited from: Sahaquiel9102 [CC BY-SA 3.0 (https://creativecommons.org/licenses/by-sa/3.0)].

References

  1. Vascular plant morphology. Topic 11, Parenchyma. Recovered from biologia.edu.ar.
  2. Ground tissue / fundamental tissue. Recovered from usepn.org.
  3. Mechanical or support fabrics. Collenchyma. Gramma University. Recovered from udg.co.cu.
  4. R. Moore, D. Clark, K.R. Stern (1998). Botany. William C Brown Pub. 832 pp.
  5. A. M. Gonzalez. Plant tissues: Meristems and fundamental system. Hypertexts in the area of ​​biology. Recovered from biologia.edu.ar.
  6. Vegetable Tissues. Atlas of plant and animal histology. Recovered from mmegias.webs.uvigo.es.
  7. Ground tissue. Wikipedia. Recovered from en.wikipedia.org.
  8. Meristematic and fundamental tissues. Recovered from iessierrasur.es.

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